Roland Pape

Associate professor

Faculty of Technology, Natural Sciences and Maritime Sciences
Department of Natural Sciences and Environmental Health
Campus Bø (1-229)
As a vegetation/landscape ecologist with a strong background in eco-climatology, I am interested in the spatio-temporal patterns and related processes within a landscape - with special emphasis on alpine environments. I am also on ResearchGate:




DOBBERT, S., PAPE, R. & LÖFFLER, J. (2022) On Growth Patterns and Mechanisms in Arctic-Alpine Shrubs. Erdkunde, 76, 199–226.

DOBBERT, S., ALBRECHT, E.C., PAPE, R. & LÖFFLER, J. (2022) Alpine Shrub Growth Follows Bimodal Patterns Across Biomes—Unexpected Environmental Controls. Communications Biology, 5, 739.

DOBBERT, S., PAPE, R. & LÖFFLER, J. (2022) The Application of Dendrometers to Alpine Dwarf Shrubs—A Case Study to Investigate Stem Growth Responses to Environmental Conditions. Biogeosciences, 19, 1933–1958.

LÖFFLER, J., ALBRECHT, E.C., DOBBERT, S., PAPE, R. & WUNDRAM, D. (2022) Dendrometer Measurements of Mediterranean-Alpine Dwarf Shrubs and Micro-Environmental Drivers of Plant Growth—Dataset from Long-Term Alpine Ecosystem Research in the Sierra Nevada, Spain (LTAER-ES). Erdkunde, DP311202.

PAPE, R. (2022) Loss of Untouched Land. - In: Tryland, M. (ed.): Arctic One Health. Challenges for Northern Animals and People, pp. 121-138.

WEHNER, A., HEIN, N., BECKERS, N., DOBBERT, S., PAPE, R. & LÖFFLER, J. (2022) Data from: Early Snow Melt and Diverging Thermal Constraints Control Body Size in Arctic-Alpine Spiders. Dryad Digital Repository

WEHNER, A., HEIN, N., BECKERS, N., DOBBERT, S., PAPE, R. & LÖFFLER, J. (2022) Early Snow Melt and Diverging Thermal Constraints Control Body Size in Arctic-Alpine Spiders. Biological Journal of the Linnean Society, 138, 1–13.


DOBBERT, S., PAPE, R., LÖFFLER, J. (2021) How does Spatial Heterogeneity Affect Inter- and Intraspecific Growth Patterns in Tundra Shrubs? Journal of Ecology, 109, 4115–4131.

DOBBERT, S., PAPE, R., LÖFFLER, J. (2021) A New Mechanistic Understanding of Ecophysiological Patterns in a Widespread Alpine Dwarf Shrub – Refining Climate-Growth Relationships. Biogeosciences (Pre-Print).

DOBBERT, S., PAPE, R., LÖFFLER, J. (2021) Contrasting Growth Response of Evergreen and Deciduous Arctic-Alpine Shrub Species to Climate Variability. Ecosphere, 12(8), e03688.

DOBBERT, S., PAPE, R., LÖFFLER, J. (2021) Continent-Wide Modelling of Potential Habitats and Connectivity for the Eurasian Lynx (Lynx lynx) in Europe. Erdkunde, 75, 69–85.

LÖFFLER, J., DOBBERT, S., PAPE, R., WUNDRAM, D. (2021) Dendrometer Measurements of Arctic-Alpine Dwarf Shrubs and Micro-Environmental Drivers of Plant Growth---Dataset from Long-Term Alpine Ecosystem Research in Central Norway. Erdkunde, DP311201.


SANDHAGE-HOFMANN, A., LÖFFLER, J., KOTZE, E., WEIJERS, S., WINGATE, V., WUNDRAM, D., WEIHERMÜLLER, L., PAPE, R., DU PREEZ, C. C. & AMELUNG, W. (2020) Woody Encroachment and Related Soil Properties in Different Tenure-Based Management Systems of Semiarid Rangelands. Geoderma, 372, 114399.

LÖFFLER, J. & PAPE, R. (2020) Thermal Niche Predictors of Alpine Plant Species. Ecology, 101, e02891. Doi:10.1002/ecy.2891


HEIN, N., PÉTILLON, J., PAPE, R., FEILHAUER, H., VANSELOW, K. A. & LÖFFLER, J. (2019) Broad-Scale Rather than Fine-Scale Environmental Variation Drives Body Size in a Wandering Predator (Araneae, Lycosidae). Arctic, Antarctic, and Alpine Research, 51, 315–326.

FRINDTE, K., PAPE, R., WERNER, K., LÖFFLER, J. & KNIEF, C. (2019) Temperature and Soil Moisture Control Microbial Community Composition in an Arctic-Alpine Ecosystem along Elevational and Micro-Topographic Gradients. The ISME Journal. Doi: 10.1038/s41396-019-0409-9.


ZIMMERMANN, S., DUBOVYK, O., OLDENBURG, C., PAPE, R. & LÖFFLER, J. (2018) Lichen Cover Mapping in Southern Norway—An Analysis with Remote Sensing and GIS. GIS Science, 2, 6071.

WEIJERS, S., PAPE, R., LÖFFLER, J. & MYERS-SMITH, I. H. (2018) Contrasting Shrub Species Respond to Early Summer Temperatures Leading to Correspondence of Shrub Growth Patterns. Environmental Research Letters. Doi: 10.1088/1748-9326/aaa5b8


PAPE, R. & LÖFFLER, J. (2017) Determinants of Arctic-Alpine Pasture Resources—The Need for a Spatially and Functionally Fine-Scaled Perspective. Geografiska Annaler: Series A, Physical Geography, 99, 353370. Doi: 10.1080/04353676.2017.1368833


PAPE, R. & LÖFFLER, J. (2016) Towards a Process-Based Biogeography of Reindeer: Scaling Space, Time, and Organizational Levels of Space Use. Norwegian Journal of Geography, 70, 230246. Doi: 10.1080/00291951.2016.1203018

PAPE, R. & LÖFFLER, J. (2016) Broad-Scale Assumptions on Available Pasture Resources and Reindeer´s Habitat Preferences Shown to be Decoupled from Ecological Reality of Arctic-Alpine Landscapes. Erdkunde, 70, 169192.

PAPE, R. & LÖFFLER, J. (2016) Spatial Patterns of Alpine Phytomass, Primary Productivity, and related Calorific Resources. Ecosphere, 7 (6), e01347. Doi:10.1002/ecs2.1347.


PAPE, R. & LÖFFLER, J. (2015) Seasonality of Habitat Selection Shown to Buffer Alpine Reindeer Pastoralism Against Climate Variability. Ecosphere, 6 (12), 260. Doi: 10.1890/ES15-00169.1

PAPE, R. & LÖFFLER, J. (2015) Ecological Dynamics in Habitat Selection of Reindeer—An Interplay of Spatial Scale, Time, and Individual Animal´s Choice. Polar Biology, 38, 18911903.Doi: 10.1007/s00300-015-1750-8


HEIN, N., PAPE, R., FINCH, O.-D. & LÖFFLER, J. (2014) Alpine Activity Patterns of Mitopus morio (Fabricius, 1779) are Induced by Variations in Temperature and Humidity at Different Scales in Central Norway. Journal of Mountain Science, 11, 644655. Doi: 10.1007/s11629-013-2913-0


PAPE, R. & LÖFFLER, J. (2012) Climate Change, Land Use Conflicts, Predation and Ecological Degradation as Challenges for Reindeer Husbandry in Northern Europe—What Do We Really Know After Half a Century of Research? Ambio, 41, 421434. Doi: 10.1007/s13280-012-0257-6.


LÖFFLER, J., ANSCHLAG, K., BAKER, B., FINCH, O.-D., DIEKKRÜGER, B., WUNDRAM, D., SCHRÖDER, B., PAPE, R. & LUNDBERG, A. (2011) Mountain Ecosystem Response to Global Change. Erdkunde, 65, 191215. Doi: 10.3112/erdkunde.2011.02.06


WUNDRAM, D., PAPE, R. & LÖFFLER, J. (2010) Alpine Soil Temperature Variability at Multiple Scales. Arctic, Antarctic, and Alpine Research, 42, 117128. Doi: 10.1657/1938-4246-42.1.117


PAPE, R., WUNDRAM, D. & LÖFFLER, J. (2009) Modelling Near-Surface Temperature Conditions in High Mountain Environments—An Appraisal. Climate Research, 39, 99109. Doi: 10.3354/cr00795


BÄR, A., PAPE, R., BRÄUNING, A. & LÖFFLER, J. (2008) Growth-Ring Variations of Dwarf Shrubs Reflect Regional Climate Signals in Alpine Environments rather than Micro-Climatic Differences. Journal of Biogeography, 35, 625636. Doi: 10.1111/j.1365-2699.2007.01804.x

FINCH, O.-D., LÖFFLER, J. & PAPE, R. (2008) Assessing the Sensitivity of Melanoplus frigidus(Boheman, 1846) (Orthoptera: Acrididae) to Different Weather Conditions: A Modelling Approach Focussing on Development Times. Insect Science, 15, 167178. Doi: 10.1111/j.1744-7917.2008.00198.x

LÖFFLER, J. & PAPE, R. (2008) Diversity Patterns in Relation to the Environment in Alpine Tundra Ecosystems of Northern Norway. Arctic, Antarctic, and Alpine Research, 40, 373381. Doi: 10.1657/1523-0430(06-097) 


LÖFFLER, J., PAPE, R. & WUNDRAM, D. (2006) The Climatologic Significance of Topography, Altitude and Region in High Mountains—A Survey of Oceanic-Continental Differentiations of the Scandes. Erdkunde, 60, 1524. Doi: 10.3112/erdkunde.2006.01.02


PAPE, R. & LÖFFLER, J. (2004) Spatio-Temporal Near-Surface Temperature Variation in High Mountain Landscapes. Ecological Modelling, 178, 483501. Doi: 10.1016/j.ecolmodel.2004.02.019

LÖFFLER, J. & PAPE, R. (2004) Across Scale Temperature Modelling Using a Simple Approach for the Characterisation of High Mountain Ecosystem Complexity. Erdkunde, 58, 331348. Doi: 10.3112/erdkunde.2004.04.04

LÖFFLER, J., LUNDBERG, A., RÖSSLER, O., BRÄUNING, A., JUNG, G., PAPE, R. & WUNDRAM, D. (2004) The Alpine Treeline under Changing Land Use and Changing Climate: Approach and Preliminary Results from Continental Norway. Norwegian Journal of Geography, 58, 183193. Doi: 10.1080/00291950410002421